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Early-Middle Pleistocene transitions: the land-ocean evidence
Conference report
Martin J. Head
Department of Geography
University of Cambridge
A one-day international conference on the “Early-Middle Pleistocene transitions:
the land-ocean evidence” was held in the Department of Geography, University
of Cambridge on April 4, 2003. The scientific programme comprised nine
invited talks and six posters. The meeting was sponsored by the Godwin
Institute for Quaternary Research, INQUA Commission on Stratigraphy and INQUA
Subcommission for European Quaternary Stratigraphy, The Quaternary Research
Association, and the ICS Subcommission on Quaternary Stratigraphy.
It was organised by Dr Martin J. Head and Dr Philip L. Gibbard. About
50 participants from England, France, Germany, Italy, Norway, Russia, and
The Netherlands made this a truly international event.
The transition from Early to Middle Pleistocene is marked
by fundamental changes in Earth’s climatic cyclicity. Orbital obliquity
at 41 ka cycles which had dominated the earlier part of the Pleistocene was
superceded progressively about a million years ago by a 100 ka rhythm of
climate change and increased-amplitude climatic oscillations. This
change has become known as the Mid-Pleistocene Revolution. The glacial–interglacial
world in which we now live is the result of these changes, and the impact
on terrestrial and marine biota has been profound and manifold. However,
the timing and correlation of events during this transition has been problematic,
leading to uncertainties over cause and effect. The purpose of this
meeting was to explore the biotic responses to climatic and physical changes
that characterized the Early–Middle Pleistocene transition in both the marine
and terrestrial realms. In doing so we hoped to explore the very origins
of our present biota.
The first talk, by Mark Maslin (UCL, London), addressed
the Mid-Pleistocene Revolution from an Earth orbital perspective and questioned
the assumption that eccentricity was responsible for 100 ka cyclicity in
the Middle Pleistocene. Disabused of this notion, we were told that
the more likely driving force was non-linear responses to orbital obliquity
paced by every fouth or fifth precessional cycle. Precession is therefore
the key to understanding 100 ka cyclicity in the Middle Pleistocene.
And precession makes sense to some paleontologists who regard seasonality
as a key to understanding their records.
Helga Flesche (“Kikki”) Kleiven (Bergen) talked about
her research on deep-sea cores in the northern and southern hemispheres that
is aimed at assessing climate change at sub-orbital frequencies across the
E–M Pleistocene transition. She showed that these sub-orbital changes, which
are well known for the past 500 ky, are constantly present throughout the
E–M Pleistocene transition. Furthermore, her work has revealed a similar
pattern of millennial-scale climate oscillations at both the northern and
southern hemispheres, implying a link in climate variability between both
hemispheres.
Erin McClymont (Durham) continued the marine theme by
looking at the E–M Pleistocene transition from a lipid biomarker perspective.
Her research, based on deep-sea cores in the North and South Atlantic and
equatorial Pacific, has allowed a comparison of high- and low-latitude climate
change at a resolution of about 5 ky. Results show important fluctuations
in carbon inputs to the ocean floor that lead climate changes as observed
in published records.
Enno Schefuß (Bremen) went on to discuss his own
research on lipid biomarkers from the equatorial Atlantic. Long-term
equatorial warming during the transition seems to be explained by a progressive
reduction in Atlantic thermohaline circulation that would have led to weakening
heat export from the tropical oceans to high latitudes. Although tropical
sea surface temperature was found to closely match the pattern of high-latitude
ice-volume changes, tropical warming at the end of glacial periods occurred
significantly earlier than decreases in global ice volume. This underscores
the role of the tropics in amplifying global climate changes.
Jean-Pierre Suc (Lyon) took us away from the deep-sea
realm and into world of Mediterranean pollen analysis, where 14 localities
allow the documentation of Late Pliocene to Middle Pleistocene climate cycles.
Filtering out the influences of latitude and altitude, an understanding of
the temporal changes in vegetational development is now emerging. In
southern Italy, for example, interglacials of the earliest Plio-Pleistocene
are signaled by Taxodiaceae, whereas those of the earliest Middle Pleistocene
are characterized by deciduous oak. The need continues, however, to
perform parallel analyses of oxygen isotopes and pollen on coastal marine
sediments.
After lunch, we were treated to two talks on human evolution
and adaptation across the E–M Pleistocene transition. The first, by
John McNabb (Southampton), discussed the spread of hominins from Africa into
Europe. The importance was stressed of understanding why it was not
until the middle of the Middle Pleistocene in Europe that a strong hominin
signal emerges. The second talk, by Mike Petraglia (Cambridge), shed
new light on hominid adaptations by reviewing current field investigations
in the Arabian Peninsula and Indian Subcontinent. Mike concluded that
the transition marks an increasing ability of hominids to transcend environmental
controls through greater behavioural flexibility and innovation.
The final talk, by Thijs van Kolfschoten (Leiden), dealt
with the European record of mammals across the transition. Notable features
are the End-Villafranchian “Event” representing a faunal turnover that is
mainly caused by the migration of a number of larger mammals, and the evolutionary
radiation of middle sized voles (e.g. Microtus). We were shown that
a number of Microtus species have sufficiently wide geographic ranges to
facilitate correlation between western and eastern Europe and even with Asia
across the Urals. The stratigraphic utility of these fossils should
therefore be taken into account when deciding where formally to place the
boundary between Early and Middle Pleistocene.
The presentations also included a report of the ICS working
group on the Early–Middle Pleistocene boundary (compiled by Davide Castradori
and presented by Martin Head) and a round-table discussion on the boundary
chaired by Charles Turner (Open University & Cambridge).
A poster session ran throughout the day and comprised
presentations on the vegetational history from the Crotone Basin in southern
Italy (L. Capraro and colleagues), the loess record of northern Eurasia (Andrey
Dodonov), orbital and suborbital variability in North Atlantic deep-water
circulation (P. Ferretti and colleagues), aminostratigraphy and integrated
land-sea correlations in the North Sea basin (two posters by T. Meijer and
P. Cleveringa), and floral evolution of the Colombian Andes (Torres and Hooghiemstra).
The conference provided an excellent overview of research
developments from both marine and terrestrial realms. It is perhaps
not greatly overstated to regard the E–M Pleistocene transition as one of
the major ‘events’ in Earth history, and the high quality of all the presentations
did justice to its importance. A volume on the E–M Pleistocene transition
is being prepared and will include many of the presentations at this conference.
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